An Excursion through my herbarium

              1 Cuad. &>t. Canar. 28: 53-63 (1916) 1917
I l I
An Excursion Through my nerbarium
by G. Kunkel "
(recibido en la redaccih: 10.2.1977)
Durante una revisión de material de herbario encontré varias
formas críticas que se dan a conocer en el texto a continuación. Se
describe 8 nuevas formas o variedades y se propone 16 nuevas com-binaciones
nomenclatóricas. Se cita, además, 4 adiciones para la
flora canaria en general, y 8 taxones como adiciones insulares.
While revising herbarium material the author describes
8 new forms or varieties of vascular plants from the Canary
Islands. For 16 taxa or subtaxa new combinations are also
proposed. Furthermore, 4 species are cited for the first time
P,, +L., P,,,,,, Tml,-A.l, ,-A Q m-nn;n, q w n l n n q l ~rlrlitinn Wi- IUI b l l c b a l i a i y lalaliun, aiiu u aprLlLa ULL LVLUL U ~ ~ L Y L U A -
thin the archipelago.
ASTER ACEAE - - -
Centaurea: Cheirolophus. The genus Cheirolophus Cass. is
recognized as being segregated from Centaurea L. (viz. Flora
Europaea 4, l976), and most corresponding Canary plants ha-ve
been dealt with by J. Holub (1973, 1974). However, the
following taxa remain to be adapted:
Cheirolophus arbutifolius (Svent.) Kunkel, comb. nov.
Centaurea arbutifolia Sventenius, Bol.1nst.Nac.Invest.
Agron. 22 (125) : 7 (1950),
an endemic shrub from Gran Canaria, and
Cheiroluphus cana~ieizsis (Willd. ) Holub
ssp. subexpinnata (Burch.) Kunkel, comb. et stat. nov.
Centaurea canariensis Willd. var. subexpinnata Bur-
* Camino Viejo 15, TAFIRA ALTA, Las Palmas de Gran Canaria.
chard, Bibl.Bot. H.98: 211 (1929).
m1 ri_ii_a_ geügi-apiiicaiiy weii isoiaied and morphoiogicaiiy
most distinctive subspecies is endemic in the Teno re-gion
of Tenerife.
Desérving further recognition
Cheirolophus ghomerythus (Svent. ) Holub
var. integrifolius (Svent.) Kunkel, comb. nov.
Centaurea ghomerytha Svent. var. integrifolia., Ad-dit.
Flor. Canar. p.73 (1960);
Cheirolophus junonianus (Svent. ) Holub
var. isoplexiphyllus (Svent.) Kunkel, comb. nov.
Centaurea junoniana Svent. var. isoplexiphylla
Svent,. Bol. 1nst.Nac.Invest.Agron. 15 (79) : 179
(1946);
Cheirolop hus satarataensis (Svent. ) Holub
ssp. dariasi (Svent.) Kunkel, comb. nov.
Centaurea satarataensis Svent. ssp. dariasi Svent.,
Addit. Flor. Canar. p. 73 (1960), and
Cheirolophus svcntcnii (A. Santos) Kunkel, comb. nov.
Centaurea sventenii A.Santos, Vieraea 4: 232 (1974)
1975, another endemic species, only recently described
from ¿a Paima (Canary isiands j. - The author thanks
Dr. J. Holub for his valuable indications concerning the
Cheirolophus species endemic in these islands.
Tolpis glabrescens Kammer
This species, only recently described from Tenerife (Ana-gaj,
seems to have a second locality in the Canary Islands:
cliffs of Guayedra below Tamadaba (Kunkel 17424: 27-VIII-
1974). Although 1 have not seen original material from Te-nerife,
Dr. Kammer's description fits our material very well;
however further comparative investigation is required.
BORAGINACEAE
Ec hium bonnetii Coincy
New to La Gomera: Kunkel 17974 (31-111-1975), La La-
ja, 400 m. The species was already known from Tenerife,
Gran Canaria, and (subspecies) Fuerteventura.
Tolpis cf. glahresccns Kimrner. Material
from Gran Canaria (Guaycdra).
BRASSICACEAE
Erysimum scoparium (Brouss. ex Willd.) Wettst.
Osterrr. Bot. Ztschr. 39: 283 (1889)
var. lindleyi (Webb ex Christ) Kunkel, comb. nov.
Cheiranthus (Dichroanthus) scoparius Willd. var.
lindleyi Webb ex Christ
Engl. Bot. Jahrb. 9: 89 (1888).
mi- - __ ..- . -L-I
U ~ :v d l i e ~ yw as descri'ued by Christ U. c. j as beionging
to Cheiranthus, a fact obviously overlooked by Mendoza-
Heuer (1972).
Lobularia intermedia Webb & Berth.
var. elongata Pitard
This seems to be a recognizabie variety, with narrow
leaves which are algo longer than usual; the infructescense
is very dense. In Gran Canaria this variety is known to occur
near to Ayacata, 1.300 m., and near to the Aserradero, 1.250
m.
MuiLhioia boiieana '%eM ex Christ
This endemic of the eastern Canary Islands was also
found (Kunkel 13187) in La Graciosa. Contrary to the opi-nion
of P. Sunding (1971) 1 believe this to be a "good spe-cies",
to be kept separate from the more freauent M. fruticu-losa.
- The aiithor thankr A . Hansen (C~pnhagen) fer
dealing with several specimens of this genus.
Matthiola fruticulosa (L.) Maire
This species ,which is frequent in parts of Lanzarote, in
its s.str. is an addition to our list of Canary plants. My mate-rial
frcm Pi'ertc Sxdinu 2nd Puerte las Nieves (U1312 C u ~ a -
ria) probably belongs to the same species. - Near to Tahi-che
(Lanzarote) we found specimens of large plants with a
woody base arid woody lower branches, forming cushions up
to 30 cm. in diameter. It remains to decide which subspecific
name may apply to our material, as M. bolleana (s.str.) is con-sidered
to be distinct and endemic in the eastern islands.
CAMPANULACEAE
Canarina canariensis (L.) Vatke, Linnaea 38: 700 (1874)
var. angustifolia Kunkel, var. nov.
A forma typica foliis angustioribus (1 - 1,5 cm. lat.) lan-ceolatis
et floribus rninoribus differt.
Holotypus: Kunkel 14557 (G) 10-1-1972, Tenerife, Seno
350 m.
Fi i r t .h~rd i f f~r enwisn 'he shape 2nd margin nf the se-pals
seem to distinguish the Gran Canarian plants from ma-terial
as found in Tenerife, for example.
CARYOPHYLLACEAE
H ~ f ~-r- .nv vrvh- .ivf nfn-n. -nv -f.n- - -nvvo cii (J. Gay) GraebE. & $/Iattf.
Contrary to the opinion as expressed by Chaudhri (1968)
1 suggest that Helerochiton has a good reason to be kept as
a gerius, segregated from Herniaria where the species con-cerned
are treated on subspecific level.
The material from Fuerteventura (Jandía, i.e. Kunkel
12586), because of the more fragile pale branches, its sprea-ding
habit, and the longer internodes may be recognized as
ssp. bollei (F. Hermann) Kunkel, comb. et stat. nov.
Herniaria bollei F. Hermann, Fedd. Repert. 42: 221
(1937).
Heterochiton canariensis (Chaudhri) Kunkel, comb. nov.
Herniaria canaricnsis Chaudhri, Rev. Paronychiinae,
Utrecht, p.383 (1968).
The species is supposed to be endemic in Tenerife. 1 have
not seen material of Herniaria pujosii Sauv. & Vindt. H.fru-ticma
L., H.e~icifolicS OK~ESH.., mu~~i t a i , iMt>ci~-h . m FT.urw-bica
Hand.-Mazz. but they should, probably, be treated
equally whereas H.FLernistemon J. Gay in Duch. seems to be
a true Herniaria.
Minuartia we.b bi.i McNeill & Bramwell TL, ., ,,.,-,,,A t, 11, ,-A,-;, 4- W , . ~ ~ t ~ . ~ ~ ~ t ~ ~ L l l C J y c L L c J LJ ouyyuocu b u uc C l I U L l L L L L 111 1 ULl LiL v L I I b U L u.
Its citation for Tenerife (Eriksson et al., 1974) seems erro-neous.
However it was recently discovered in Lanzarote whe-re
it grows in the Riscos de Guinate, at 350 m a.s.1.
Polycarpaea nivea (Ait. ) Webb, Specil. Gorg. p.104 (1849)
f. quadraiigixlare Kunkel, forma nova
Suffrutex decurnbens, argenteus, frondosus; folia decus-sata
emittens aspectus quadrangularis.
Holotypus : Kunkel 17409 (G) , 17-N-1974, Gran Cana-ria,
Playa de Jinámar, 20 m., rare.
This smd ! rn-l ounuf wi th v. -"-A v g - j densdgr and de~i-~.ssa1tp~. -
ves should not be confused with var. diffusa of Pitard.
Polscarpaea nivea (Ait.) Webb
var.longifolia Kunkel, var. nov.
Suffr.u t.e x compactus usque ad 40 c m alt.; lamina carno- sci, mí-de-glabm, oblanc zdc t~W S ~ Z : CG W 3,V l m vY n et 7
mm. Lata.
Holotypus: Kunkel 18972 (G), 30-111-1976, Fuerteven-tura,
Barranco ias Cabras, ¿u m. Rare.
Po1 ycarpaea smithii Link
mIr1 lis species is supp"sed i" lDe eii&iiiic in La T>a!na. It
has been cited for Hierro, without a locality. IIowever it
grows also in La Gomera (Kunkel 11470), in walls of the
Barranco La Laja, at 600 m. approximstely.
Sclerocephalus arabicus Boiss.
N curious record, obviousiy new io the Lanary ísiands:
Fuerteventura, Jandía 150 cm, on rocky slope (Kunkel15098)
The plants are small, probably annual, 8 to 10 cm. tall; the
glaucous and linear leaves are up to 1 cm long, and the fruits
are globose-glomerate and spiny.-The species was (in my
herharii'im) mixed-i'ip with P t ~ r n tnh .?!..~f r, n m t h s~am P l n r a -
lity.
CISTACEAE
Cistus symphytifolius Lam., Encycl. Méth. 2: 15 (1786)
ssp. symphytifolius
ssp. leucophyllus (Spach) Kunkel, comb. et stat. nuv.
Rhodocistus berthelotianus Spach var. leucophyllus
Spach, Hist. Vég. Phan. 6: 93 (1838)
Cistus symphytifolius Lam. var. leucophyllus (Spach)
Dansereau (1939).
This subspecies is endemic to Gran Canaria where the
area of both subspecies are well defined and no hybridization
is known to occur.
Helianthemum canariense (Jacq.) Pers., Syn. P1. 2: 78 (1806)
var subglabratum Kunkel, var. nov.
A var. canariense foliis subglabris differt. Suffrutex den-se
ramificatis usque ad 40 cm. alt.
Holotypus: Kunkel 12721 (G), 11-IV-1969, Gran Cana-ria,
La Calzadalsan Lorenzo, 250 m.
This more shrubby, darker leaved and almost glabrous
variety grows on dry slopes, together with specimens of the
iy pica1 val ieiy .
Helianthemum thymiphy llum Sventenius
After publishing a second locality (Roque Aceituno, near
to Femez: Kunkel 1976), a new locality of this obviously
long overlooked species may be added: Caldera Blanca, NW
oi limaniaya, where Andryaia gianciuiosa, iviicromeria vu-ria,
Asplenitim hemionitis and Adiantum capillus-veneris al-so
occur.
CONVOLVULACEAE
co?.vgglvwlus flnyi,clur L. fil., Sijppl. Pi., p. 136 (1781).
var. angustif olius (Pit. ) Kunkel, comb. nov.
Rhodorhiza florida Webb & Berth. var. angustifolia
Pitard, Fedd. Repert. 9: 379 (1311).
This variety was described by Pitard (1. c.) as Rhodor-hiza,
a fact obviously overlooked by Mendoza-Heuer (1971),
and the revalidation of the variety is herewith suggested.
Ipomoea hederacea (L.) Jacq.
This ornamental annual climber, much cultivated in
Gran Canaria, Lanzarote, etc., was found invading neighbou-ring
gardens in severa1 places. Our record (Kunkel 15859)
from Lanzarote (San Antonio gardens) might be listed as an
addition to the aliens of the Canary flora. The material was
named by A. Hansen (Copenhagen).
CRASSULACEAE
Aeonium percarneum (Murr.) Pitard in Pit. & Pr., Iles Ca-nar.
p. 191 (1908).
var. guiaense Kunkel, var. nov.
A typo (var. percarneo) habito minore densiore, foliis
Aichryson laxum (Haw.) Bramwell, Bol. Inst. Nac. Invest.
Agron. 59: 207 (1968)
f. subglabrum Kunkel, forma nova
Herba biennis subglabris usque ad 80 cm altis, foliis lon-gipetiolatis,
Lamina usque ad 6 cm. lat.
Holotypus: Kunkel 18485 (G), 12-VII-1975, Gran Cana-ria,
Vaisendero 75U m.
This might be (?) an adaptive form to an extremely
moist and more shaded environment. Almost a hundred spe-cimens
of this form were observed, over severa1 years.
Arbutus unedo L., the Mediterranean "Madroño"
In the vicinity of San Mateo (Gran Canaria), near a loca-lity
known as "El Montañón"), we found an admirable stand
of older trees of this species, freely flowering and fruiting.
However no natural regeneraiion has beeri noiiced.
LAURACEAE
Laurus azorica (Seub.) Franco var. longifolia (0. Ktze.) Kun-kel
'Trees of this iong-ieaved and poor-Iiowered variety nave
been found in Tenerife, on the old foot-path from the Forest
House towards Taganana.
PERIPLOCACEAE
Periploca laevigata Ait., Hort. Kew. p.YO1 i1789)
f. undulata Kunkel, forma nova
A forma typica lamina crenato-undulata dif f ert. Foliis
lanceolatis usque ad 9 cm longis.
Holotypus : Kunkel 15191 (G) , 1.-VIII-1972, Gran Ca-naria,
Barranco de Arguineguín, 600 m. Rare.
f. obovata Kunkel, forma nova
Differl a forma typica foliis obovatis, lamina 4-5 cm
longa et 2,5-3 cm latu.
Holotypus: Kunkel 13993a (G), 26-111-1971, Gran Ca-naria,
Andén Verde towards Tirma 400 m.; rare.
PLANTAGINACEAE
Psyllium afrum (L.) Mirb.
This species, usuallg an annual, is quite common in the
Canary Islands, especially in the eastern group of the archipe-lago.
However, certain material from Lobos Island, Alegran-za
and La Graciosa has larger, elongated flower heads very
similar to Psyllium arenarium (Waldst. & Kit.) Mirb.
The three remainin species of this genus are endemic in
the Macaronesian islands and are listed as follows:
Psyllium arborescens (Poir.) Mirb., Hist. Pl., ed. 2: 333 (1830)
ssp. arborescens
A rather polymorphic species of the central and western
;"lc,,-.A" ; m n l , l A ; n m IiJILIIIUi>, I I I L I U U I I I ~
var. canescens (Svent.) Kunkel, comb. nov.
Plantago arborescens Poir. var. canescens Sventenius:
Addit. Flor. Canar. (Madrid) p.62 (1960), and
var. mascaensis (Svent.) Kunkel, comb. nov.
Plantago arborescens Poir. var. mascaensis Svente-nius,
Addit. Flor. Canar. (Madrid), p.61 (1960).
both of which are endemic in Tenerife. In a wider sense the
Madeiran form should be cited as
ssp. maderensis (Decne.) Kunkel, comb. et stat. nov.
Plantago maderensis Decne. in DC., Prodr. 13 (1) :
733 (1852)
P.arborescens ssp. maderensis (Decne.) Hansen &
Kunkel (1972)
Psyllium famarae (Svent.) Kunkel, comb. nov.
Plantago famarae Sventenius, Addit. Flor. Canar.
(Madrid) p.b3 (1960): a species endemic in the north-wes'tern
part of Lanzarote, and
Psyllium webbii (Barn.) Soják, Cas. Národ. Muz. Ser. Nalur.
(Praha) 140: 131; 1972. A rare endemic of higher re-
gions of Gran Canaria, Tenerife, and La Palma.
SAPOTACEAE
Argania spinosa (L.) Skeels
In the Barranco de Agaete (Gran Canaria), at about 400
m. a.s.l., exists a sizeable specimen of this African plant, ob-viously
introduced but well established and which may
spread just as other introduced plants have.
SOLANACEAE
Sn- l..n- .n. ..- 7- i... m . .j nsm.in.oides Paxt.
This species is abundant near to the Forest Kouse in the
Taganana (Anaga) range, in Tenerife, where it grows on
r-"a&sides aiid invades tlie foresti tse1f =hile &-bing inla .+
re1 trees.
Kcicrcnces
CBAUDiHRI, M.N., 1968: A revision of tlie Paronyclziinae. - Diss Utrecht, 440 pp.
ERIKSSOU. O., HANSEN, A. & P SUNDING, 1974: ,Flora of Macaronesia.
Check-lisr of vascular plants 1974. - Umed University, 66 pp.
HOLUB, J., 1973: Some new nomenclatura1 combinations in Centaureinx (Aste-raceae).
- Preslia 45: 142-146.
IíVLüB, I., 1773: Somc nc*r íívmcnc!ut;ru! c~m?iii,utio~isn Cectaureitiw (Ate-raceae)
11. - Preslia 46: 225-229.
KAMMER, F., 1976: Tolpis ghbrescens Kammer, eine neue Kompositen-Art aus
Macaronesien - Rot Jalzrb. Syst. 97: 155-160.
KUNKEL, G., 1976: Enunleración de las plantas vasculares del Parque Nacional
de Timanfaya (Lanzarote), con notas adicionales. - Cuad. Bot. Canar.
26/27: 41-58.
MENDOZA-HEUFR, 1, 1971: Aportación al conocimiento del género Convol-vulu~
s en la zona Macaronésica - Cuad. Bot. Cannr. 12: 22-34.
MENDOZA-HEUER, 1 , 1972: Acerca del género Erysimum (Cruciferae) en la
zona Macaronésica. - Cuad. Bot. Canar. 14/15: 17-26.
SUNDING, P., 1971: Additions to the vdaculdr iloia o£ Lanzarote and Fuerteven-tura.
- Cuad. Bot. Canar. 13: 15-20.